Liverwort

Anemone hepatica (syn. Hepatica nobilis), the common hepatica, liverwort, liverleaf, kidneywort, or pennywort, is a species of flowering plant in the buttercup family Ranunculaceae, native to woodland in temperate regions of the Northern Hemisphere. This herbaceous perennial grows from a rhizome.

Description

Anemone hepatica grows 5–15 cm (2–6 in) high. Leaves and flowers emerge directly from the rhizome, not from a stem above ground.

The leaves have three lobes and are fleshy and hairless, 7–9 cm (2+3⁄4–3+1⁄2 in) wide and 5–6 cm (2–2+1⁄4 in) long. The upper side is dark green with whitish stripes and the lower side is violet or reddish brown. Leaves emerge during or after flowering and remain green through winter.

The flowers are blue, purple, pink, or white and appear in winter or spring. They have five to ten oval showy sepals and three green bracts.

Taxonomy

The taxonomy of the genus Anemone and its species is not fully resolved, but phylogenetic studies of many species of Anemone and related genera indicate that species of the genus Hepatica should be included under Anemone because of similarities both in molecular attributes and other shared morphologies. The circumscription of the taxon is also debated, some authors listing the North American var. acuta and var. obtusa, while other list them as the separate species A. acutiloba and A. americana, respectively.

Varieties

Varieties of Anemone hepatica that are sometimes recognized include:

• Anemone hepatica var. japonica, a synonym of Hepatica nobilis var. japonica Nakai, is native to the Russian Far East, China, Korea, and Japan
• Anemone hepatica var. acuta, a synonym of Hepatica acutiloba DC., is native to eastern North America
• Anemone hepatica var. obtusa, a synonym of Hepatica americana (DC.) Ker Gawl., is native to eastern North America

Distribution and habitat

It is found in woods, thickets and meadows, especially in the mountains of continental Europe, North America and Japan.

Ecology

Hepatica flowers produce pollen but no nectar. In North America, the flowers first attract Lasioglossum sweat bees and small carpenter bees looking in vain for nectar. Then when the stamens begin to release pollen, the bees return to collect and feed on pollen. Mining bees sometimes visit the flowers, but prefer flowers that produce both nectar and pollen.

Toxicity

Like other Ranunculaceae, fresh liverwort contains protoanemonin and is therefore slightly toxic. By drying the herb, protoanemonin is dimerized to the non-toxic anemonin.

Uses

Medieval herbalists believed it could be used to treat liver diseases, and may be used in current folk medicine.

Under the name Hepatica nobilis, which is regarded as a synonym, this plant has gained the Royal Horticultural Society's Award of Garden Merit.

Culture

It is the official flower of the Sweden Democrats political party in Swedish politics.

Gallery

References

• Pignatti, S. (1982). Flora d'Italia. Vol. 1. Edagricole. p. 277.

External links

• "Hepatica nobilis (Liverleaf)". BioLib.cz. Retrieved 15 January 2021.
• Flora of Pennsylvania
• Anemone hepatica in the CalPhotos photo database, University of California, Berkeley